Spring Boot WebMvcTest, ¿qué está causando IllegalArgumentException aquí?

Tengo el siguiente WebMvcTest en un proyecto Spring Boot. Tengo habilitada la autenticación de token HTTPS y JWT. La prueba funcionó antes de que tuviera HTTPS y JWT, pero después de agregarlos da una IllegalArgumentException.

Por mi vida no puedo entender de dónde viene la excepción. ¿Alguien puede señalarme en la dirección correcta?

Clase de prueba:

@RunWith(SpringRunner.class)
@WebMvcTest(controllers = ActionController.class)
public class ActionControllerTests {

    @Autowired
    private MockMvc mvc;

    @MockBean
    private ActionService service;

    @MockBean
    private UserDetailsService userDetailsService;

    @MockBean
    private JwtRequestFilter jwtRequestFilter;

    @Test
    @WithMockUser
    public void testGetActions_returns_result_from_service() throws Exception {
        int actionId = 1;
        Action action = new Action();
        action.setId(actionId);
        List<Action> actionsList = Arrays.asList(action);

        given(service.getActions()).willReturn(actionsList);

        mvc.perform(get("/actions").secure(true)
                .contentType(MediaType.APPLICATION_JSON))
                .andExpect(status().isOk())
                .andExpect(jsonPath("$", hasSize(1)))
                .andExpect(jsonPath("$[0].id", Matchers.is(actionId)));
    }
}

Lanza esta IllegalArgumentException:

java.lang.IllegalArgumentException: Attribute name must not be null

    at org.springframework.util.Assert.notNull(Assert.java:198)
    at org.springframework.mock.web.MockHttpServletRequest.setAttribute(MockHttpServletRequest.java:756)
    at javax.servlet.ServletRequestWrapper.setAttribute(ServletRequestWrapper.java:257)
    at javax.servlet.ServletRequestWrapper.setAttribute(ServletRequestWrapper.java:257)
    at org.springframework.web.filter.OncePerRequestFilter.doFilter(OncePerRequestFilter.java:117)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.security.web.authentication.logout.LogoutFilter.doFilter(LogoutFilter.java:116)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.web.filter.CorsFilter.doFilterInternal(CorsFilter.java:92)
    at org.springframework.web.filter.OncePerRequestFilter.doFilter(OncePerRequestFilter.java:119)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.security.web.header.HeaderWriterFilter.doHeadersAfter(HeaderWriterFilter.java:92)
    at org.springframework.security.web.header.HeaderWriterFilter.doFilterInternal(HeaderWriterFilter.java:77)
    at org.springframework.web.filter.OncePerRequestFilter.doFilter(OncePerRequestFilter.java:119)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.security.web.context.SecurityContextPersistenceFilter.doFilter(SecurityContextPersistenceFilter.java:105)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.security.web.context.request.async.WebAsyncManagerIntegrationFilter.doFilterInternal(WebAsyncManagerIntegrationFilter.java:56)
    at org.springframework.web.filter.OncePerRequestFilter.doFilter(OncePerRequestFilter.java:119)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.security.web.access.channel.ChannelProcessingFilter.doFilter(ChannelProcessingFilter.java:157)
    at org.springframework.security.web.FilterChainProxy$VirtualFilterChain.doFilter(FilterChainProxy.java:334)
    at org.springframework.security.web.FilterChainProxy.doFilterInternal(FilterChainProxy.java:215)
    at org.springframework.security.web.FilterChainProxy.doFilter(FilterChainProxy.java:178)
    at org.springframework.mock.web.MockFilterChain.doFilter(MockFilterChain.java:134)
    at org.springframework.web.filter.RequestContextFilter.doFilterInternal(RequestContextFilter.java:100)
    at org.springframework.web.filter.OncePerRequestFilter.doFilter(OncePerRequestFilter.java:119)
    at org.springframework.mock.web.MockFilterChain.doFilter(MockFilterChain.java:134)
    at org.springframework.web.filter.FormContentFilter.doFilterInternal(FormContentFilter.java:93)
    at org.springframework.web.filter.OncePerRequestFilter.doFilter(OncePerRequestFilter.java:119)
    at org.springframework.mock.web.MockFilterChain.doFilter(MockFilterChain.java:134)
    at org.springframework.test.web.servlet.MockMvc.perform(MockMvc.java:183)
    at nl.kars.lms.controller.ActionControllerTests.testGetActions_returns_result_from_service(ActionControllerTests.java:57)
    at java.base/jdk.internal.reflect.NativeMethodAccessorImpl.invoke0(Native Method)
    at java.base/jdk.internal.reflect.NativeMethodAccessorImpl.invoke(NativeMethodAccessorImpl.java:62)
    at java.base/jdk.internal.reflect.DelegatingMethodAccessorImpl.invoke(DelegatingMethodAccessorImpl.java:43)
    at java.base/java.lang.reflect.Method.invoke(Method.java:566)
    at org.junit.runners.model.FrameworkMethod$1.runReflectiveCall(FrameworkMethod.java:50)
    at org.junit.internal.runners.model.ReflectiveCallable.run(ReflectiveCallable.java:12)
    at org.junit.runners.model.FrameworkMethod.invokeExplosively(FrameworkMethod.java:47)
    at org.junit.internal.runners.statements.InvokeMethod.evaluate(InvokeMethod.java:17)
    at org.springframework.test.context.junit4.statements.RunBeforeTestExecutionCallbacks.evaluate(RunBeforeTestExecutionCallbacks.java:74)
    at org.springframework.test.context.junit4.statements.RunAfterTestExecutionCallbacks.evaluate(RunAfterTestExecutionCallbacks.java:84)
    at org.springframework.test.context.junit4.statements.RunBeforeTestMethodCallbacks.evaluate(RunBeforeTestMethodCallbacks.java:75)
    at org.springframework.test.context.junit4.statements.RunAfterTestMethodCallbacks.evaluate(RunAfterTestMethodCallbacks.java:86)
    at org.springframework.test.context.junit4.statements.SpringRepeat.evaluate(SpringRepeat.java:84)
    at org.junit.runners.ParentRunner.runLeaf(ParentRunner.java:325)
    at org.springframework.test.context.junit4.SpringJUnit4ClassRunner.runChild(SpringJUnit4ClassRunner.java:251)
    at org.springframework.test.context.junit4.SpringJUnit4ClassRunner.runChild(SpringJUnit4ClassRunner.java:97)
    at org.junit.runners.ParentRunner$3.run(ParentRunner.java:290)
    at org.junit.runners.ParentRunner$1.schedule(ParentRunner.java:71)
    at org.junit.runners.ParentRunner.runChildren(ParentRunner.java:288)
    at org.junit.runners.ParentRunner.access$000(ParentRunner.java:58)
    at org.junit.runners.ParentRunner$2.evaluate(ParentRunner.java:268)
    at org.springframework.test.context.junit4.statements.RunBeforeTestClassCallbacks.evaluate(RunBeforeTestClassCallbacks.java:61)
    at org.springframework.test.context.junit4.statements.RunAfterTestClassCallbacks.evaluate(RunAfterTestClassCallbacks.java:70)
    at org.junit.runners.ParentRunner.run(ParentRunner.java:363)
    at org.springframework.test.context.junit4.SpringJUnit4ClassRunner.run(SpringJUnit4ClassRunner.java:190)
    at org.junit.runner.JUnitCore.run(JUnitCore.java:137)
    at com.intellij.junit4.JUnit4IdeaTestRunner.startRunnerWithArgs(JUnit4IdeaTestRunner.java:68)
    at com.intellij.rt.junit.IdeaTestRunner$Repeater.startRunnerWithArgs(IdeaTestRunner.java:33)
    at com.intellij.rt.junit.JUnitStarter.prepareStreamsAndStart(JUnitStarter.java:230)
    at com.intellij.rt.junit.JUnitStarter.main(JUnitStarter.java:58)

Gracias un montón.